J. G. M. Thewissen
Scientific Bibliography (excluding abstracts)
Last updated, July 2007

NOTE: Some PDF's are password protected for copyright reasons. Please email thewisse@neoucom.edu for access.

Books and Monographs

In press. Thewissen, J. G. M. and S. Nummela (editors). Senses on the Threshold, the Evolution of Sense Organs in Secondarily Aquatic Tetrapods. University of California Press.

2002. Perrin W. P., B. W. Würsig, and J. G. M. Thewissen (editors). Encyclopedia of Marine Mammals. 1414 pages. Academic Press. Second Edition in preparation (Elsevier).

1998. Thewissen, J. G. M. (editor). The Emergence of Whales, Evolutionary Patterns in the Origin of Cetacea. 475 pages. Plenum Press (now Kluwer).

1996. Thewissen, J.G.M., S. I. Madar, and S. T. Hussain. Ambulocetus natans, an Eocene cetacean (Mammalia) from Pakistan. Courier Forschungs-Institut Senckenberg, 190:1-86.

1990. Thewissen, J.G.M. Evolution of Paleocene and Eocene Phenacodontidae (Mammalia, Condylarthra). Papers on Paleontology, University of Michigan, 29:1-107.

Articles and Chapters

2007. Nummela, S., J.G.M. Thewissen, S. Bajpai, T. Hussain, and K. Kumar. Sound transmission in archaic and modern whales: anatomical adaptations for underwater hearing. Anatomical Record 290:716-733. Download PDF

2007. Thewissen, J. G. M. and J. Heyning. Chapter 15, Embryogenesis and Development in Stenella attenuata and other Cetaceans. Pp. 305-327 in: Reproductive Biology and Phylogeny of Cetacea. D. Miller. (ed.) Science Publishers, Inc., Enfield, NH. Download PDF

2007. Thewissen, J. G. M., and M. A. Taylor. Aquatic adaptations in Amniotes. Pp. 310-322. In: B. K. Hall (ed.), ]Fins into Limbs, Evolution, Development, and Transformation. Univ. Chicago Press. Download PDF

2006. Bajpai, S., V. V. Kapur, J. G. M. Thewissen, D. Das, B. N. Tiwari. New early Eocene cambaythere (Perissodactyla, Mammalia) from the Vastan Lignite Mine (Gujarat, India) and an evaluation of cambaythere relations. Journal of the Palaeontological Society of India 51-101-110. Download PDF

2006. Bajpai, S., Thewissen, J. G. M., V. V. Kapur, B. N. Tiwari, and Sahni, A. Eocene and Oligocene sirenians (Mammalia) from Kachchh, India. Journal of Vertebrate Paleontology 26:400-410. Download PDF (password required)

2006. Nummela, S.,S. T. Hussain, and J. G. M. Thewissen. Cranial anatomy of Pakicetidae (Cetacea, Mammalia). Journal of Vertebrate Paleontology. 26:746-759. Download PDF (password required)

2006. Thewissen, J. G. M., Cohn, M. J., Stevens, L. S., Bajpai, S., Heyning, W. E., and Horton, W. E. Jr. Developmental basis for hind limb loss in dolphins and the origin of the cetacean bodyplan. Proceedings of the National Academy of Sciences, U.S.A.. 103:8414-8418.2006. Download PDF

2005. Bajpai, S., A. V. V. Kapur, J. G. M. Thewissen, D. P. Das, B. N. Tiwari, R. Sharma, and N. Saravanan. Early Eocene primates from Vastan Lignite Mine, Gujarat, Western India. Journal of the Palaeontological Society of India, 50. Download PDF

2005. Bajpai, S., V. V. Kapur, J. G. M. Thewissen, B. N. Tiwari, and D. P. Das. First fossil marsupials from India: Early Eocene Indodelphis n. gen and Jaegeria n. gen. from Vastan Lignite Mine, District Surat, Gujarat. Journal of the Palaeontological Society of India 50:147-151. Download PDF

2005. Nishihara, Y. Satta, M. Nikaido, J.G.M. Thewissen,, M. J. Stanhope, and N. Okada. A retroposon analysis of Afrotherian Phylogeny. Molecular Biology and Evolution 22:1823-1833. Download PDF

2005. Plikus, M. V., M. Zeichner-David, J.-A.Mayer, J. Reyna, P. Bringas, J. G. M. Thewissen, M. L. Snead, Y. Chai, and C.-M. Chuong. Morphoregulation of teeth: modulating the number, size, shape, and differentiation by tuning Bmp activity. Evolution and Development 7:440-457. Download PDF

2005. Thewissen, J. G. M., S. Bajpai, and S. T. Hussain. New insectivorous placentals from the Eocene of Pakistan. Journal of the Palaeontological Society of India, 50:37-41. Download PDF

2004. Nummela, S., Kosove, J. E., Lancaster, T., and Thewissen, J. G. M. Variation in lateral mandibular wall thickness in Tursiops truncatus and other odontocetes. Marine Mammal Science 20:491-497. Download PDF (password required)

2004. Nummela, S., J. G. M. Thewissen, S. Bajpai, S. T. Hussain, and K. Kumar. Eocene evolution of whale hearing. Nature 430:776-778. Download PDF

2003. Rage, J.-C. , S. Bajpai, J. G. M. Thewissen, and B. N. Tiwari. Early Eocene snakes from Kutch, Western India, with a review of the Palaeophiidae. Geodiversitas 25:645-716. Download PDF

2002. Kumar, K., M. W. Hamrick, and J. G. M. Thewissen. Middle Eocene prosimian primate from the Subathu Group of Kalakot, northwestern Himalaya, India. Current Science (New Delhi) 79:1478-1482. Download PDF

2002. Thewissen, J. G. M., and E. M. Williams. The early evolution of Cetacea (whales, dolphins, and porpoises). Annual Review of Ecology and Systematics 33:73-90. Download PDF

2002. Bajpai, S, and J. G. M. Thewissen. Vertebrate fauna from Panandhro Lignite Field (Lower Eocene), District Kachchh, western India. Current Science 82:507-509. Download PDF

2002, Madar, S.I., J. G. M. Thewissen, and S. T. Hussain. Additional holotype remains of Ambulocetus natans (Cetacea, Ambulocetidae), and their implications for locomotion in early whales. Journal of Vertebrate Paleontology 22:405-422.

2002, Spoor, F., S. Bajpai, S. T. Hussain, K. Kumar, and J. G. M. Thewissen. Vestibular evidence for the evolution of aquatic behaviour in early cetaceans. Nature 417:163-166. Download PDF

2001, Thewissen, J. G. M., and S. Bajpai. Whale origins as posterchild for macroevolution. BioScience 5:1037-1049. Download PDF

2001, Thewissen, J. G. M., E. M. Williams, L.J. Roe, and S.T. Hussain. Skeletons of terrestrial cetaceans and the relationship of whales to artiodactyls. Nature, 413:277-281. Download PDF

2001, Maas, M.C., J.G.M. Thewissen, S. Sen, N. Kazanci, J. Kappelman. Enigmatic new ungulates from the early middle Eocene of Central Anatolia, Turkey. Journal of Vertebrate Paleontology 21:578-590. Download PDF (password required)

2001. Thewissen, J.G.M., and E.M. Williams, and S.T. Hussain. Eocene mammal faunas from northern Indo-Pakistan. Journal of Vertebrate Paleontology, 21:347-366.

2001. Maas, M. C., S. T. Hussain, J.J.M. Leinders, and J.G.M. Thewissen. A new isectolophid tapiromorph (Perissodactyla,Mammalia) from the early Eocene of Pakistan. Journal of Paleontology 75:407-417.

2001. Kinkel, M., J. G. M. Thewissen, and H. A. Oelschläger. Rotation of middle ear ossicles during cetacean development. Journal of Morphology, 249:126-131.

2001. Thewissen, J. G. M., and S. Bajpai. Dental morphology of the Remingtonocetidae (Cetacea, Mammalia). Journal of Paleontology 75:463-465.

2001. Thewissen, J.G.M. and E. L. Simons. Skull of Megalohyrax eocaenus (Hyracoidea, Mammalia) from the Oligocene of Egypt. Journal of Vertebrate Paleontology 21:98-106.

2000. S. Bajpai, and J. G. M. Thewissen. A new, diminuitive whale from Kachchh (Gujarat, India) and its implications for locomotor evolution of cetaceans. Current Science (New Delhi) 79:1478-1482. Download PDF

2000. Sánchez-Villagra, M. R., R. J. Burnham, D. C. Campbell, R.M. Feldmann, E.S. Gaffney, R.F. Kay, R. Lozsán, R. Purdy, and J.G.M. Thewissen. A new near-shore marine fauna and flora from the early Neogene of northwestern Venezuela. Journal of Paleontology74:957-968.

2000. Thewissen, J.G.M., and S.T. Hussain. Attockicetus praecursor, a new remingtonocetid cetacean from marine Eocene sediments of Pakistan. Journal of Mammalian Evolution 7:133-146.

1999. Thewissen, J.G.M., and S. I. Madar. Ankle morphology of the earliest cetaceans and its implications for the phylogenetic relations among ungulates. Systematic Biology, 48:21-30.

1998. Thewissen, J.G.M., S.T. Hussain, M. Arif, A. Aslan, S.I. Madar, and L.J. Roe. New localities of Eocene vertebrates in northern Pakistan and their significance for the origin of modern orders of mammals. Proceedings of the third GEOSAS Workshop held at Islamabad, Pakistan, March 1-5, 1998, pp. 19-34.

1998. Thewissen, J.G.M., S. I. Madar, and S. T. Hussain. Whale ankles and phylogenetic relations. Nature, 395:452.

1998. Bajpai, S., and J. G. M. Thewissen. Middle Eocene cetaceans from the Harudi and Subathu Formations of India. In: Thewissen, J. G. M., Editor: The Emergence of Whales, Evolutionary Patterns in the Origin of Cetacea. Pp. 213-233. Plenum Press.

1998. Roe, L. J., J. G. M. Thewissen, J. Quade, J. R. O'Neil, S. Bajpai, A. Sahni, and S. T. Hussain. Isotopic approaches to understanding the terrestrial to marine transition of the earliest cetaceans. In: Thewissen, J. G. M., Editor: The Emergence of Whales, Evolutionary Patterns in the Origin of Cetacea. Pp. 399-421. Plenum Press.

1998. Thewissen, J. G. M. Cetacean origins: evolutionary turmoil during the invasion of the oceans. In: Thewissen, J. G. M., Editor: The Emergence of Whales, Evolutionary Patterns in the Origin of Cetacea. 451-464. Plenum Press.

1998. Kalmey, J. K., J.G.M. Thewissen, D. E. Dluzen. Age-related size reduction of foramina in the cribriform plate. Anatomical Record 251:326-329.

1998. Maas, M.C., J. G. M. Thewissen, and J. Kappelman. Hypsamasia seni (Mammalia: Embrithopoda) and other mammals from the Eocene Kartal Formation of Turkey. Bulletin of the Carnegie Museum 34:286-297. Download PDF

1998. Thewissen, J. G. M., and S. T. Hussain. Systematic review of the Pakicetidae, Early and middle Eocene Cetacea (Mammalia) from Pakistan and India. Bulletin of the Carnegie Museum 34:220-238.

1997. Milinkovitch, M. C., and J. G. M. Thewissen. Eventoed fingerprints on cetacean ancestry. Nature, 388:622-624.

1997. Thewissen, J. G. M., and F. E. Fish. Locomotor evolution in the earliest cetaceans: functional model, modern analogues, and paleontological evidence. Paleobiology 23:482-490.

1997. Thewissen, J.G.M., and S.T. Hussain. New Kohatius (Omomyidae) from the Eocene of Pakistan. Journal of Human Evolution 32:473-477.

1997. Thewissen, J.G.M., S. I. Madar, E. Ganz, S. T. Hussain, M. Arif, H. Hussain. Fossil yak (Bos grunniens: Artiodactyla, Mammalia) from the Himalayas of Pakistan. Kirtlandia (Cleveland) 50:11-16.

1996. Aslan, A., and J. G. M. Thewissen. Preliminary evaluation of Kuldana paleosols and implications for interpreting vertebrate fossil assemblages, Kuldana Formation, Northern Pakistan. Palaeovertebrata 25:261-277. Download PDF

1996. Bajpai, S., J. G. M. Thewissen, and A. Sahni. Indocetus (Cetacea, Mammalia) endocasts from Kachchh (India). Journal of Vertebrate Paleontology, 16:582-584.

1996. Thewissen, J.G.M., L. J. Roe, J. R. O'Neil, S. T. Hussain, A. Sahni, and S. Bajpai. Evolution of cetacean osmoregulation. Nature 381:379-380. Download PDF

1995. Maas, M.C. and J. G. M. Thewissen. Enamel microstructure of Pakicetus (Mammalia: Archaeoceti). Journal of Paleontology, 69:1154-1163.

1995. Thewissen, J.G.M., and S. A. Etnier. Adhesive devices of the thumb of vespertilionoid bats (Chiroptera). Journal of Mammalogy, 76:925-936.

1995. Thewissen, J.G.M. Phylogenetic aspects of cetacean origins: a morphologic perspective. Journal of Mammalian Evolution 2:157-184.

1994. Savage, R.J.G., D.P. Domning, and J.G.M. Thewissen. Fossil Sirenia of the west Atlantic and Carribean Region. V. The most primitive known sirenian, Prorastomus sirenoides Owen, 1855. Journal of Vertebrate Paleontology 14:427-449.

1994. Thewissen, J.G.M., S.T. Hussain, and M. Arif. Fossil evidence for the origin of aquatic locomotion in archaeocete whales. Science 263:210-212.

1993. Thewissen, J.G.M. Eocene marine mammals from the Himalayan foothills. Research and Exploration, National Geographic Society, 9:125-127. Erratum: Research and Exploration 9:487.

1993. Thewissen, J.G.M., and S.K. Babcock. The implications of the propatagial muscles of flying and gliding mammals for archontan systematics. In: Primates and their Relatives in Phylogenetic Perspective (R.D.E. MacPhee, editor), Plenum, New York, pp. 91-110.

1993. Thewissen, J.G.M., and S.T. Hussain. Origin of underwater hearing in whales. Nature 361:444-445.

1992. Kay, R.F., J.G.M. Thewissen, and A.D. Yoder. The cranial anatomy of Ignacius graybullianus and the affinities of the Plesiadapiformes. American Journal of Physical Anthropology 89:477-498.

1992. Thewissen, J.G.M. Temporal data in phylogenetic systematics: An example from the mammalian fossil record. Journal of Paleontology, 66:1-8.

1992. Thewissen, J.G.M., and S.K. Babcock. The origin of flight in bats: To go where no mammal has gone before. BioScience 42:340-345.

1992. Thewissen, J.G.M., and D.P. Domning. The role of phenacodontids in the origin of the modern orders of ungulate mammals. Journal of Vertebrate Paleontology 12:494-504.

1992. Thewissen, J.G.M., and M.C. McKenna. Paleobiogeography of Indo-Pakistan: A response to Briggs, Patterson, and Owen. Systematic Biology 41:248-251. Download PDF

1991. Thewissen, J.G.M. Limb osteology and function of the primitive Paleocene ungulate Pleuraspidotherium, with notes on Tricuspiodon and Dissacus (Mammalia). Geobios (Lyon) 24:483-495.

1991. Thewissen, J.G.M., and S.K. Babcock. Distinctive cranial and cervical innervation of wing muscles: New evidence for bat monophyly. Science 251:934-936.

1990. Thewissen, J.G.M. Comment on "Paleontological view of the ages of the Deccan Traps, the Cretaceous/Tertiary boundary, and the India-Asia collision." Geology 18:185, 187-188. Download PDF

1990. Thewissen, J.G.M., and S.T. Hussain. Postcranial osteology of the most primitive artiodactyl, Diacodexis pakistanensis (Dichobunidae). Anatomia, Histologia, Embryologia (Berlin), 19:37-48.

1989. Thewissen, J.G.M. Mammalian frontal diploic vein and the human foramen caecum. Anatomical Record 223:242-244.

1989. Thewissen, J.G.M., and P.D. Gingerich. Skull and endocranial cast of Eoryctes melanus, a new palaeoryctid (Mammalia: Insectivora) from the early Eocene of Western North America. Journal of Vertebrate Paleontology 9:459-470.

1988. Shoshani, J., C.A. Goldman, and J.G.M. Thewissen. Orycteropus afer. Mammalian Species, 300:1-8. Download PDF

1987. Thewissen, J.G.M., and J.-L. Franzen. Fossillagerstaette Messel. Cranium (Utrecht), 4:86-90.

1987. Thewissen, J.G.M., and P.D. Gingerich. Revision of Probathyopsis (Mammalia, Dinocerata) from the late Paleocene and early Eocene of western North America. Contributions from the Museum of Paleontology, University of Michigan, 27:195-219.

1987. Thewissen, J.G.M., P.D. Gingerich, and D.E. Russell. Artiodactyla and Perissodactyla (Mammalia) from the early-middle Eocene Kuldana Formation of Kohat (Pakistan). Contributions from the Museum of Paleontology, University of Michigan, 27:247-274.

1987. Thewissen, J.G.M., and G.R. Smith. Vespertilionid bats (Chiroptera, Mammalia) from the Pliocene of Idaho. Contributions from the Museum of Paleontology, University of Michigan, 27:237-245.

1986. Thewissen, J.G.M., and D.M. Badoux. The descriptive and functional myology of the fore-limb of the Aardvark (Orycteropus afer Pallas, 1766). Anatomischer Anzeiger, (Jena) 162:109-123. Download PDF

1985. Thewissen, J.G.M. Cephalic evidence for the affinities of Tubulidentata. Mammalia, 49:257-284.

1983. Hussain, S.T., P.Y. Sondaar, S.M.I. Shah, J.G.M. Thewissen, E.F.H.M. Cousin, and C.F. Spoor. Fossil mammal bones of Pakistan, a field atlas. Part I: the artiodactyl astragalus. Memoirs of the Geological Survey of Pakistan, 14:4-30.

1983. Russell, D.E., J.G.M. Thewissen, and D. Sigogneau-Russell. A new artiodactyl (Mammalia) from the Eocene of North-West Pakistan. Part II: Cranial osteology. Proceedings of the Koninklijke Nederlandse Akademie voor Wetenschappen, Series B, 86:285-300.

1983. Thewissen, J.G.M., D.E. Russell, P.D. Gingerich, and S.T. Hussain. A new artiodactyl (Mammalia) from the Eocene of North-West Pakistan. Dentition and classification. Proceedings of the Koninklijke Nederlandse Akademie voor Wetenschappen, Series B, 86:153-180.